Click
here to close Hello! We notice that
you are using Internet Explorer, which is not supported by Echinobase
and may cause the site to display incorrectly. We suggest using a
current version of Chrome,
FireFox,
or Safari.
Proc Biol Sci
2018 Jan 31;2851871:. doi: 10.1098/rspb.2017.2590.
Show Gene links
Show Anatomy links
Whole-body photoreceptor networks are independent of ''lenses'' in brittle stars.
Sumner-Rooney L
,
Rahman IA
,
Sigwart JD
,
Ullrich-Lüter E
.
Abstract
Photoreception and vision are fundamental aspects of animal sensory biology and ecology, but important gaps remain in our understanding of these processes in many species. The colour-changing brittle star Ophiocoma wendtii is iconic in vision research, speculatively possessing a unique whole-body visual system that incorporates information from nerve bundles underlying thousands of crystalline ''microlenses''. The hypothesis that these might form a sophisticated compound eye-like system regulated by chromatophores has been extensively reiterated, with investigations into biomimetic optics and similar supposedly ''visual'' structures in living and fossil taxa. However, no photoreceptors or visual behaviours have ever been identified. We present the first evidence of photoreceptor networks in three Ophiocoma species, both with and without microlenses and colour-changing behaviour. High-resolution microscopy, immunohistochemistry and synchrotron tomography demonstrate that putative photoreceptors cover the animals'' oral, lateral and aboral surfaces, but are absent at the hypothesized focal points of the microlenses. The structural optics of these crystal ''lenses'' are an exaptation and do not fulfil any apparent visual role. This contradicts previous studies, yet the photoreceptor network in Ophiocoma appears even more widespread than previously anticipated, both taxonomically and anatomically.
Aizenberg,
Calcitic microlenses as part of the photoreceptor system in brittlestars.
2001, Pubmed,
Echinobase
Aizenberg,
Calcitic microlenses as part of the photoreceptor system in brittlestars.
2001,
Pubmed
,
Echinobase
Bielecki,
Ocular and extraocular expression of opsins in the rhopalium of Tripedalia cystophora (Cnidaria: Cubozoa).
2014,
Pubmed
Blevins,
Spatial vision in the echinoid genus Echinometra.
2004,
Pubmed
,
Echinobase
Bok,
Here, There and Everywhere: The Radiolar Eyes of Fan Worms (Annelida, Sabellidae).
2016,
Pubmed
Burke,
A genomic view of the sea urchin nervous system.
2006,
Pubmed
,
Echinobase
D'Aniello,
Opsin evolution in the Ambulacraria.
2015,
Pubmed
,
Echinobase
Delroisse,
De Novo Adult Transcriptomes of Two European Brittle Stars: Spotlight on Opsin-Based Photoreception.
2016,
Pubmed
,
Echinobase
Delroisse,
High opsin diversity in a non-visual infaunal brittle star.
2014,
Pubmed
,
Echinobase
Garm,
Visual navigation in starfish: first evidence for the use of vision and eyes in starfish.
2014,
Pubmed
,
Echinobase
Gorzelak,
Microlens arrays in the complex visual system of Cretaceous echinoderms.
2014,
Pubmed
,
Echinobase
Jackson,
Orientation to Objects in the Sea Urchin Strongylocentrotus purpuratus depends on apparent and not actual object size.
2011,
Pubmed
,
Echinobase
Land,
Image formation by a concave reflector in the eye of the scallop, Pecten maximus.
1965,
Pubmed
Lesser,
Sea urchin tube feet are photosensory organs that express a rhabdomeric-like opsin and PAX6.
2011,
Pubmed
,
Echinobase
Leung,
Unconventional Roles of Opsins.
2017,
Pubmed
Müller,
A cryptochrome-based photosensory system in the siliceous sponge Suberites domuncula (Demospongiae).
2010,
Pubmed
Nilsson,
Advanced optics in a jellyfish eye.
2005,
Pubmed
Nilsson,
The evolution of eyes and visually guided behaviour.
2009,
Pubmed
O'Hara,
Restructuring higher taxonomy using broad-scale phylogenomics: The living Ophiuroidea.
2017,
Pubmed
Petie,
Crown-of-thorns starfish have true image forming vision.
2016,
Pubmed
,
Echinobase
Polishchuk,
Coherently aligned nanoparticles within a biogenic single crystal: A biological prestressing strategy.
2017,
Pubmed
,
Echinobase
Raible,
Opsins and clusters of sensory G-protein-coupled receptors in the sea urchin genome.
2006,
Pubmed
,
Echinobase
Ramirez,
The Last Common Ancestor of Most Bilaterian Animals Possessed at Least Nine Opsins.
2016,
Pubmed
Ramirez,
Understanding the dermal light sense in the context of integrative photoreceptor cell biology.
2011,
Pubmed
Richter,
Invertebrate neurophylogeny: suggested terms and definitions for a neuroanatomical glossary.
2010,
Pubmed
Schindelin,
Fiji: an open-source platform for biological-image analysis.
2012,
Pubmed
Speiser,
A chiton uses aragonite lenses to form images.
2011,
Pubmed
Sumner-Rooney,
Whole-body photoreceptor networks are independent of 'lenses' in brittle stars.
2018,
Pubmed
,
Echinobase
Ullrich-Lüter,
Unique system of photoreceptors in sea urchin tube feet.
2011,
Pubmed
,
Echinobase
Vukusic,
Photonic structures in biology.
2003,
Pubmed
Whitfield,
Presumptive ciliated receptors associated with the fibrillar glands of the spines of the echinoderm Amphipholis squamata.
1983,
Pubmed
,
Echinobase
Yerramilli,
Spatial vision in the purple sea urchin Strongylocentrotus purpuratus (Echinoidea).
2010,
Pubmed
,
Echinobase