Lach R and Salamon MA (2016), Late Cretaceous crinoids (Echinodermata) from t...

ECB-ART-50019

Palaontol Z 2016 Jan 01;903:503-520. doi: 10.1007/s12542-016-0313-9.

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Late Cretaceous crinoids (Echinodermata) from the southwestern margin of the Holy Cross Mts. (southern Poland) and phylogenetic relationships among bourgueticrinids.

Lach R , Salamon MA .

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A systematic account of crinoids from the Upper Coniacian-Lower Campanian of the southwestern margin of the Holy Cross Mountains in southern Poland is presented. Seven crinoid taxa [Marsupites testudinarius (von Schlotheim), Bourgueticrinus ellipticus (Miller), Bourgueticrinus sp., I.? granosus Valette, Isocrinus? sp., Nielsenicrinus carinatus Roemer and Austinocrinus bicoronatus (von Hagenow)] are described and illustrated. The new material from Poland extends down the stratigraphic range of Austinocrinus bicoronatus to the Lower Campanian. Morphometric data support that Bourgueticrinus ellipicus and B.? suedicus are conspecific. Taphonomy and paleoecology of recorded crinoid assemblages are discussed. Phylogeny of Cretaceous bourgueticrinids is also revisited.

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	Fig. 1. a Map of Poland with enlarged studied localities (slightly modified from Remin 2004, 2010). b Section of the outcrop no. 1. and distribution of collected crinoid taxa (modified after Remin 2010; biostratigraphy after Walaszczyk 1992)
	Fig. 2. Dimensions of bourgueticrinid cups used in the present study (idea after Głuchowski 1987, modified)
	Fig. 3. Santonian crinoid Marsupites testudinarius (von Schlotheim) from the southwestern margin of the Holy Cross Mountains and GPIH. Scale bars 1 cm. a–g Thecae. Upper Santonian. Lägerdorf. GPIH 40.1114/16-19. h–j Thecal plates. Upper Santonian. Lipnik–Kije Section. GIUS 9-3651/Mt/1-3
	Fig. 4. Upper Cretaceous crinoids from the southwestern margin of the Holy Cross Mountains. Upper Santonian/Lower Campanian. All from Lipnik–Kije Section. Scale bars 1 cm. A–F. Bourgueticrinus ellipticus (Miller). A1, A2 Thecae. Specimen with high collar and sloping articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/1. B1, B2 Thecae. Specimen with high collar and sloping articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/2. C1, C2 Thecae. Specimen with small collar and sloping articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/3. D1, D2 Thecae. Specimen with high collar and sloping articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/4. E1, E2 Thecae. Specimen with high collar and sloping articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/5. F1, F2 Thecae. Specimen with small collar and sloping articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/6
	Fig. 5. Upper Cretaceous crinoids from the southwestern margin of the Holy Cross Mountains. Upper Santonian/Lower Campanian. All from Lipnik–Kije Section. Scale bars 1 cm. a–f. Bourgueticrinus ellipticus (Miller). A1, A2 Thecae. Specimen with small collar and sloping articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/7. B1, B2 Thecae. Specimen with small collar and horizontal articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/8. C1, C2 Thecae. Specimen without collar and horizontal articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/9. D1, D2 Thecae. Specimen without collar and sloping articular faces. Upper and lateral views, respectively. GIUS 9-3651/Be/10. E1–E3 Thecae. Specimen with small collar, sloping articular faces, and elliptical cup base. Upper, lateral, and lower views, respectively. GIUS 9-3651/Be/11. F1–F3 Thecae. Specimen without collar, sloping articular faces, and circular cup base. Upper, lateral, and lower views, respectively. GIUS 9-3651/Be/12
	Fig. 6. Late Cretaceous crinoids from the southwestern margin of the Holy Cross Mountains. All from Lipnik–Kije Section. Scale bars 1 mm. A–C Bourgueticrinus sp. A1, A2 Thecae. Lateral and upper views, respectively. Lower Campanian. GIUS 9-3651/B/3. B–C Columnals. Lateral view. Upper Coniacian. GIUS 9-3651/B/10-11. D Isocrinus? sp. Columnals. Articular face. Upper Coniacian. GIUS 9-3651/I/3. E–G Isocrinus? granosus Valette. E. Columnals. Articular face. Upper Coniacian. GIUS 9-3651/Ig/3. F, G Cirrals. Lateral and upper views, respectively. Upper Coniacian. GIUS 9-3651/Ig/11-12. H–K Nielsenicrinus carinatus Roemer. H–I. Columnals. Articular face and lateral views, respectively. Upper Santonian. GIUS 9-3651/Nc/1-2. J, K Brachials. Upper view. Upper Santonian. GIUS 9-3651/Nc/4-5. L Austinocrinus bicoronatus (von Hagenow). Columnals. Articular face. Lower Campanian. GIUS 9-3651/Ab/1
	Fig. 7. Relationship between the height of the collar and the total height of the cup (specimens from Geologisch-Paläontologisches Museum in Hamburg) showing strong positive correlation (Spearman's rank–order correlation coefficient P = 0.25022, p = 1.16E−05)
	Fig. 8. Relationship between the height of the collar and the cup width in the narrowest point (specimens from Geologisch-Paläontologisches Museum in Hamburg) showing strong positive correlation (Spearman's rank–order correlation coefficient P = 0.21836, p = 0.000138)
	Fig. 9. Relationship between the height of the collar and the cup width in the widest point (specimens from Geologisch-Paläontologisches Museum in Hamburg) showing strong positive correlation (Spearman's rank–order correlation coefficient P = 0.16179, p = 0.004969)
	Fig. 10. Suggested phylogeny of Cretaceous bourgueticrinid crinoids

References [+] :

Baumiller, Post-Paleozoic crinoid radiation in response to benthic predation preceded the Mesozoic marine revolution. 2010, Pubmed, Echinobase